Last updated 21 February 2026: extensively updated Introduction, listing with many new species, and Bibliography.

A look at the

Subfamily GEKKONINAE - "True" Geckos

Family GEKKONIDAE

Genus PACHYDACTYLUS - Thick-Toed or Rough Geckos

A large (about 35 species) and rather ancient African genus, being considered ancestral to some of the other gekkonid genera such as Tarentola (Branch). Members of this genus are rarely seen in the trade, partly because there is a perceived lack of interest in "brown geckos" (as Walls calls them), but mainly because South Africa and Namibia do not allow exports. The two species that were once considered common pets, P. turner and P. bibronii, are now considered to be members of Chondrodactylus instead.

The genus is characterised by the following: dilated toe tips, usually with undivided scansors: no femoral pores, but preanal pores may be present. Body scales small, granular and non-overlapping, with scattered, large keeled tubercles. Most if not all are nocturnal. They live in a diverse range of habitats and feed mainly on arthropods. The range of this genus is centred on southern Africa, with some reaching East Africa, the northernmost limit of their distribution.

In the table below, measurements are total-length approximations. They are taken mainly from the snout-vent length (SVL) given in Branch, and doubled, since the tail length of Pachydactylus species is about 50% of the total length (see SHDA). We have used centimetres rather than inches for this table due to the small size of most of the species.

There is still much work to be done on the individual species on this table, which will be carried out over a period of time.

In 2005 the two species of web-footed geckos, formerly assigned to their own genus Palmatogecko, were reassigned to Pachydactylus. Both species hail from the Namib Desert in South-West Africa and have the useful adaptation of webbing between their toes to enable them to more easily cross the sandy terrain in their habitat. The Namib Desert itself, with its fine sand (at least in the range of P. rangei), dense fogs caused by cold coastal currents and near-equanimity of day- and night temperatures (ie unlike most deserts, the temperature hardly drops at night), is not the easiest habitat to replicate in a terrarium, and for this reason it is suggested that these beautiful but rather specialised geckos should only be kept by gecko specialists and/or those with plenty of experience in recreating unusual habitats.

NOTES: SHDA refers to the Field Guide to Reptiles of East Africa, Branch to Snakes and Other Reptiles of Southern Africa. To go to the Bibliography from a given entry, click on B: to return to the Quick Index, click on I.

	QUICK INDEX
P. acuminatus	P. affinis, Transvaal Thick-Toed Gecko	P. amoenus
P. angolensis, Angola Large-Scaled Gecko, Angola Thick-Toed Gecko	P. atorquatus, Augrabie’s Gecko, Good’s Gecko	P. austeni, Austen's Thick-Toed Gecko
P. barnardi, Barnard's Thick-Toed Gecko	P. bicolor, Velvety Thick-Toed Gecko	P. bibronii, Bibron's Thick-Toed Gecko
P. boehmei	P. capensis, Cape Thick-Toed Gecko	P. caraculicus, Angolan Banded Thick-Toed Gecko
P. carinatus, Richtersveld Gecko	P. etultra, Sossus Gecko	P. fasciatus, Banded Thick-Toed Gecko
P. fitzsimonsi, Fitzsimons' Thick-Toed Gecko	P. formosus, Thick-Toed Gecko	P. gaiasensis, Brandberg Thick-Toed Gecko
P. geitje, Ocellated Thick-Toed Gecko	P. griffini	P. haackei, Haacke's Thick-Toed Gecko
P. katanganus, Katanga Thick-Toed Gecko	P. kladaroderma, Thin-Skinned Thick-Toed Gecko	P. kobosensis, Thick-Toed Gecko
P. kochii, Koch's Thick-Toed Gecko	P. labialis, Western Cape Thick-Toed Gecko	P. latirostris, Quartz Gecko
P. macrolepis, Large-Scaled Banded Gecko	P. maculatus, Spotted Thick-Toed Gecko	P. maiatoi, Maiato’s Thick-Toed Gecko
P. maraisi	P. mariquensis, Marico Thick-Toed Gecko	P. mclachani
P. monicae, Monica’s Gecko	P. montanus, Namaqua Montane Gecko, Montane Thick-Toed Gecko	P. namaquensis, Namaqua Thick-Toed Gecko
P. namibensis, Namibe Thick-Toed Gecko	P. oculatus, Golden Spotted Thick-Toed Gecko	P. oreophilus, Kaokoveld Thick-Toed Gecko
P. oshaughnessyi, O'Shaughnessy's Thick-Toed Gecko	P. otaviensis	P. ovahimba, Himba Thick-Toed Gecko
P. parascutatus	P. punctatus, Speckled Thick-Toed Gecko	P. purcelli, Purcell’s Gecko
P. rangei, Namib Sand Gecko, Namib Web-Footed Gecko	P. reconditus	P. robertsi, Large-Scaled Gecko, Shielded Thick-Toed Gecko
P. rugosus, Rough Thick-Toed Gecko	P. sansteyni, San Steyn's Thick-Toed Gecko	P. scherzi, Schertz's Thick-Toed Gecko
P. scutatus, Thick-Toed Gecko	P. serval, Thick-Toed Gecko	P. tetensis, Tete Thick-Toed Gecko
P. tigrinus, Tiger Thick-Toed Gecko	P. tsodiloensis, Tsodilo Thick-Toed Gecko	P. tuberculosus, Tuberculate Thick-Toed Gecko
P. turneri, Turner's Thick-Toed Gecko	P. vansoni, Van Son's Thick-Toed Gecko	P. vanzyli, Namib Desert Gecko
P. visseri, Visser’s Gecko	P. wahlbergii, Kalahari Ground Gecko, Wahlberg’s Kalahari Ground Gecko	P. waterbergensis
P. weberi, Weber's Thick-Toed Gecko	P. werneri, Werner’s Rough-Toed Gecko

Scientific Name	Common Name	Distribution	Size	Notes
P. acuminatus		Namibia		Formerly considered a subspecies of P. weberi.
P. affinis	Transvaal Thick-Toed Gecko	RSA (four northernmost provinces): poss. Free State & SE Botswana	6-9cm	B I
P. amoenus		RSA (Cape Province, Namaqualand)	?	For some reason this species is not mentioned in Branch. See the EMBL database listing. B I
P. angolensis	Angola Large-Scaled Gecko, Angola Thick-Toed Gecko	Angola, NW Namibia
P. atorquatus	Augrabie’s Gecko, Good’s Gecko	Republic of South Africa
P. austeni	Austen's Thick-Toed Gecko	RSA (coastal areas of W Cape to Little Namaqualand)	6-9cm	B I
P. barnardi	Barnard’s Rough Gecko, Barnard's Thick-Toed Gecko	RSA (S Little Namaqualand, Cape Province), Namibia		Formerly considered subspecies of P. rugosus. B I
P. bicolor	Velvety Thick-Toed Gecko	Namibia (NW Damaraland and Kaokoveld)	6-8cm	B I
P. bibronii	Bibron's Gecko	S. Africa, Angola, S. Tanzania	8-9"
P. boehmei		Namibia
P. capensis	Cape Thick-Toed Gecko	RSA (Cape Provinces)	8-13½cm	Description: head regularly oviform, not much broader than the neck ; snout a little longer than the diameter of the orbit. Ear-opening oval, oblique. Body depressed. Limbs moderate; digits short, rather slender, distinctly broader at the end than at the base, the dilated part with 5-6 lamellae inferiorly. Tail rather depressed, tapering, distinctly annulate. Snout covered with convex scales which are much larger than the granules of the back; hinder part of head covered with granules intermixed with large round tubercles; naso-rostrals in contact; rostral broader than high ; seven or eight upper labials; 6-7 infralabials, the anterior as large as the mental; latter about twice as long as broad, slightly narrowing posteriorly. Upper parts covered with very small granules intermixed with large, feebly keeled, roundish or suboval tubercles, arranged symmetrically on the back; abdominal scales moderate, increasing in size from throat to groin. Upper surface and sides of tail with slightly imbricate smooth scales and transverse rows of large feebly keeled tubercles; lower surface of tail with imbricate smooth scales. Coloration: light brown above, variegated with dark brown and whitish; a dark brown streak on each side of the head, passing through the eye ; lower surfaces immaculate. [SOURCE: Boulenger] B I
P. caraculicus	Angolan Banded Thick-Toed Gecko	SW Angola, Namibia (N Kaokoveld)	6-8cm	B I
P. carinatus	Richtersveld Gecko	Republic of South Africa
P. etultra	Sossus Gecko	Namibia	SVL <4.5 cm	A rupicolous species. Description: a dorsoventrally flattened species; (excluded in P. kobosensis) (Fig. 2C); snout rounded and nasal region moderately inflated; digits not long; body slender, caudal tubercles within a tail whorl well separated Scalation details: scales on snout larger than interorbital tubercles; the first supralabial narrowly enters the nostril; dorsal scalation heterogeneous, with the entire dorsum of trunk equally tuberculate. Coloration: three body bands (including nape and sacral bands) present in juveniles and adults, although sometimes obscured in the latter; bands narrow and straight-edged, lost or obscured in adults. [SOURCE: Branch et al 2011]
P. fasciatus	Banded Thick-Toed Gecko	Namibia (N Damaraland and Kaokoveld)	8-11cm	B I
P. fitzsimonsi	Fitzsimons' Thick-Toed Gecko	Namibia (Damaraland), S Angola	13-18cm	B I
P. formosus	Thick-Toed Gecko	RSA (W Cape Province)	13-18cm	Description: agrees in general with P. capensis. Differs in having the digits rather shorter and furnished with four or five lamellae inferiorly, the large dorsal tubercles strongly keeled and the small scales between them more irregular, and the ventral scales smaller. Scales on the upper part of the tail keeled. Coloration: yellowish, with brown variegations; a broad, brown, crescent-shaped band bordering the head posteriorly; four broad, brown, transverse bars on the back, broader than the interspaces between them. [SOURCE: Boulenger] B I
P. gaiasensis	Brandberg Thick-Toed Gecko	Namibia (Brandberg)	10-13cm	B I
P. geitje	Ocellated Thick-Toed Gecko	RSA (SW Cape, inc. Cape Fold mtns, Port Elizabeth and Cradock)	6-9cm	Description: snout very short, scarcely longer than the diameter of the orbit, obtuse, convex. Ear-opening subcircular. Body not much depressed. Limbs moderate; digits short, subequal, the end not much dilated, with 4-5 lamellae inferiorly. Tail: thick, rounded, tapering. Scalation details: upper surface of head, body, and limbs covered with granules of nearly equal size. Naso-rostrals generally separated; rostral scarcely broader than high; upper labials 8-9; lower labials 7-8, the anterior as long as, and broader than, the mental; latter quadrangular, not, or but little, narrowed posteriorly. Abdominal scales imbricate, equal. Caudal scales smooth, slightly imbricate, subequal, about twice as large as the granules of the back. Male with a series of 3-4 conical scales on each side of the base of the tail. Coloration: grey or brown above, with small, white, dark-edged ocelli; upper lip white; a dark streak on the side of the head and neck, passing through the eye; whitish beneath, throat dotted with brown. [SOURCE: Boulenger] B I
P. griffini		SE Namibia
P. haackei	Haacke's Thick-Toed Gecko	S Namibia	14-17cm	B I
P. katanganus	Katanga Thick-Toed Gecko	S Democratic Republic of Congo, N Malawi		Formerly considered a subspecies of P. oreophilus. Description: 6-8 thin yellow bands and larger keeled tubercles.
P. kladaroderma	Thin-Skinned Thick-Toed Gecko	RSA (inland Cape escarpment on the Nuweveldberg, through to western Karoo and on to the southern Cape fold mtns)	13-17cm	B I
P. kobosensis		Namibia	?	Very restricted range: see EMBL database listing. B I
P. kochii	Koch's Thick-Toed Gecko	Namibia (Cape Cross & vicinity)	8-11cm	Closely related to P. mariquensis. B I
P. labialis	Western Cape Thick-Toed Gecko	RSA (W Cape from Little Namaqualand to Calvinia)	6-9cm	B I
P. latirostris	Quartz Gecko	Republic of South Africa		Formerly considered a subspecies of P. mariquensis. Scalation: nasorostrals separated by granular scales: dorsal scales flattened and slightly overlapping. Namibian population is isolated.
P. macrolepis	Large-Scaled Banded Gecko	Republic of South Africa		Formerly considered a subspecies of P. mariquensis.
P. maculatus	Spotted Thick-Toed Gecko	SE Africa [[RSA (Cape Province, S. Zululand, Natal, Transkei, SE Transvall), Swaziland]	4-5"	A grey-brown gecko with dark, sharply outlined spots. It lives among stones and bush roots, in leaf litter or in holes in ground, in the latter case often sharing these with girdle-tailed lizards (Cordylid species) or scorpions. Perhaps as a result of this, many in the wild seem to have regenerated tails. Although rarely seen in the trade, in captivity it apparently tames readily. Description: agrees in size, proportions, and scutellation with P. geitje, except that the granules of the back are intermixed with scattered larger tubercles, twice or three times the size of the granules, that there are a few conical tubercles on the outer side of the tibia, and that there is one lamella less under the disks of the digits. Coloration: greyish or brownish above ; a broad dark brown streak on the side of the head, passing through the eye, converging towards its fellow on the occiput; back with four longitudinal series of large dark brown spots, those of the outer series generally, those of the inner series constantly confluent into a band; tail spotted and variegated with dark brown ; lower surfaces dotted with brown. [SOURCE: Boulenger, Branch] B I
P. maiatoi	Maiato’s Thick-Toed Gecko	Angola
P. maraisi		C Namibia
P. mariquensis	Marico Thick-Toed Gecko	RSA, Namibia	8-11cm	Description: head very convex ; snout very short, hardly as long as the diameter of the orbit, round. Ear-opening oval or subcircular. Body not much depressed. Limbs long; digits rather short, the end slightly dilated, with three lamellae inferiorly; the outer toe inserted a good deal below the fourth. Tail: cylindrical, tapering, without annuli. Scalation details: upper surface of head, body, and limbs covered with granules of subequal size; dorsal scales granular and abutting; naso-rostrals in contact; rostral broader than high; 7-8 upper labials; 6-7 lower labials, gradually decreasing in size, the anterior as long as, and broader than, the mental, which is very slightly narrowed posteriorly. Abdominal scales equal. Caudal scales equal, smooth, slightly imbricated, much larger than the granules of the back, Male with a series of four conical scales on each side of the base of the tail. Coloration: grey above, with reddish-brown blackish-margined markings ; these markings are a spot on the nose, another on the forehead, a semicircular broad bar surrounding the back of the head, stretching from eye to eye, and posteriorly directed angular broad bars on the back and tail; lower surfaces whitish, immaculate. [SOURCE: Boulenger] B I
P. mclachani		Namibia
P. monicae	Monica’s Gecko	Namibia, Republic of South Africa
P. montanus	Namaqua Mountain Gecko, Montane Thick-Toed Gecko	Namaqua, Republic of South Africa	?
P. namaquensis	Namaqua Thick-Toed Gecko	Namaqualand: poss. Namibia	14-17cm	B I
P. namibensis	Namibe Thick-Toed Gecko	Angola
P. oculatus	Golden Spotted Thick-Toed Gecko	Southern Africa	7-11cm	White ventral and grey-brown dorsal surfaces with dark band running along both sides and dark stripes across the back as far as the tail. A lateral dark band extends across the eye to the snout. B I
P. oreophilus	Kaokoveld Thick-Toed Gecko	Namibia (Brandberg), S Angola	8-11cm	B I
P. oshaughnessyi	O'Shaughnessy's Thick-Toed Gecko	N Zimbabwe, W Mozambique, Malawi, Zambia	9-11cm	Description: head regularly oviform, not much broader than the neck; snout obtuse, subtriangular, a little longer than the diameter of the orbit. Ear-opening very small, oval, oblique. Limbs moderate; digits short, of subequal width throughout their length, with 4-5 lamella; inferiorly. Body not much depressed. Tail: rounded, very thick in its anterior half, ending in a thin point. Scalation details: snout covered with convex scales which are much larger than the granules of the back; the remainder of the head covered with very small granules intermixed with larger ones; naso-rostrals in contact; rostral broader than high; 8 upper labials; 7 lower labials, first as long as, and broader than, the mental; latter a little narrowed posteriorly, twice as long as broad. Back covered with small granules intermixed with large, keeled, conical tubercles arranged rather irregularly. Outer side of femur and tibia with conical tubercles. Throat granulate, belly covered with moderate-sized imbricated scales. Tail: covered with rather large, equal, strongly imbricate, cycloid, smooth scales. Coloration: head yellowish, bordered by a blackish streak extending from one nostril to the other, passing through the eye and conturning the occiput; this streak bordered posteriorly by a broader cream-coloured band; 3-4 yellow crossbands on back (4-5 in Zambia); back and upper surface of tail light brown, the former with two, the latter with seven, large, transverse, cream coloured, blackish- edged spots; limbs and lower surfaces uniform cream-coloured. [SOURCE: Boulenger]
P. otaviensis		Namibia
P. ovahimba	Himba Thick-Toed Gecko	Angola
P. parascutatus		N Namibia	Max SVL approx <4 cm	Description: body sub-cylindrical, not depressed; most dorsal scales large, rounded, strongly keeled, juxtaposed or weakly imbricate, in approximately 14 rows; ventral scales subimbricate, scales on dorsum of forelimb and granules between larger dorsal scales unkeeled; nostril surrounded by two postnasals, supranasal, rostral and first supralabial; enlarged scansorial lamellae under digit IV of pes 4-5. Coloration: adult dorsal pattern with numerous dark brown markings, without distinct white nuchal collar; original tail more-or-less banded; venter buff with light brown speckling. [SOURCE: Bauer et al]
P. punctatus	Speckled Thick-Toed Gecko	N Botswana, Zimbabwe, Namibia, Angola, Congo	5-7cm	Differs from P. geitje in having the snout somewhat longer and more pointed, the granules of the back and the scales of the tail larger, and the scales on the snout three or four times as large as those on the back of the head. Seven upper and six lower labials. Brown above, spotted with blackish brown; behind the eye a yellow band, blackish-edged above; lower surfaces pure white. [SOURCE: Boulenger] B I
P. purcelli		RSA (W Little Karoo), SE Namibia		Formerly considered a subspecies of P. serval. Scalation: dorsal scales granular, lacks tubercles.
P. rangei	Web-Footed Sand Gecko	S Angola, Namibia, Republic of South Africa	4"	A rather delicate but beautiful gecko with large eyes and, most distinctively, webbing between the toes of both front and back limbs. This adaptation is obviously useful for helping it to cross the fine sand of the Namib dunes. In addition to the webbing, the toes pads have adhesive lamellae in the same manner as other members of the Gekkoninae, eg Tokay Geckos, Gekko gecko. The limbs themselves seem to be most frequently described as "spindly", perhaps as a weight-bearing adaptation in addition to the webbed feet. These lizards obtain moisture by the 'capture' of dew that condenses on their scales. They are nocturnally active throughout the year, sheltering from the heat during the day in deep burrows that they usually dig themselves. Like many geckos, P. rangei can vocalise, in this case clicking and squeaking either when displaying or protesting (ie if handled). Both Mattison and Bartlett and Bartlett deal with the care of the Web-Footed Gecko: suffice it to say that the habitat requirements are fairly exact, similar in some ways to those for the popular Teratoscincus geckos. The authors reckon however that if these are met then these geckos are fairly hardy in captivity and have been bred. In appearance the Web-Footed Gecko is easily distinguished by its overall pink dorsal colour with a variety of stripes, its huge eyes with their brown-red irises and vertical pupils, and its slender form.
P. reconditus		Namibia
P. robertsi	Large-Scaled Gecko, Shielded Thick-Toed Gecko	Namibia
P. rugosus	Rough Thick-Toed Gecko	RSA [S. Little Namaqualand, Cape Province], Namibia	9-13cm	Very occasionally seen. Two former three subspecies (barnardi and formosus) are now full species. Preferred habitat is dry riverbeds and/or rocky outcrops, where they may either conceal themselves in cracks or in leaf litter, etc. Description: head oviform, very distinct from neck; snout moderate, obtuse, a little longer than the diameter of the orbit. Ear-opening oval, oblique. Body not much depressed. Limbs rather long; digits short, of subequal width throughout their length, with 4-5 lamellae inferiorly. Tail (reproduced?): short, thick. Head covered with conical tubercles intermixed with small granules; naso-rostrals separated by a granule ; rostral a little broader than high ; 10 upper labials; 9 lower labials, gradually decreasing in size; mental subtriangular. Upper parts with small granules of irregular size, intermixed with large, conical, spinose tubercles arranged irregularly. Scales on lower surfaces granular, subconical. Coloration: head brownish white, with a large, crescent-shaped, greyish-brown mark extending from eye to eye; back greyish brown, with four transverse, dentated, whitish bars, narrower than the intervals; a whitish band from the mouth to the anterior cross bars; lower surfaces whitish, speckled with brown. [SOURCE: Boulenger] B I
P. scherzi	Schertz's Thick-Toed Gecko	Namibia (coastal plain and inland escarpment, from the Brandberg north to the Kaokoveld)	5-7cm	B I
P. sansteyni	San Steyn's Thick-Toed Gecko	Namibia (Kuidas, S Kaokoveld, mouth of Cunene River)	7-9cm	B I
P. scutatus	Large-Scaled Thick-Toed Gecko	N Nambia and S Angola	6-8cm	B I
P. serval	Western Spotted Thick-Toed Gecko	RSA (S Karoo to Lower Orange River and Great Namaqualand: absent from W Cape and Little Namaqualand): S Namibia	7-9cm	B I
P. s. serval		S Namibia		Scalation: dorsal tubercles scattered and enlarged, not keeled or raised.
P. s. onscepensis		RSA (lower Orange River valley)		Scalation: dorsal tubercles scattered, raised and keeled.
P. tetensis	Tete Thick-Toed Gecko	SE Tanzania, Mozambique, Lower Zambezi valley	Max TL 18cm: avg TL 13-15cm	B I
P. tigrinus	Tiger Gecko	Zimbabwe, Republic of South Africa (N. Transvaal), E. Botswana, W. Mozambique (S. Blinkwater/Brak River)	?	A brown gecko with white stripes in a tiger pattern. I have seen this gecko offered for sale on a business web site, but other than that have no information about it.
P. tsodiloensis	Tsodilo Thick-Toed Gecko	N Botswana (Tsodilo Hills)	9-12cm	B I
P. tuberculosus	Tuberculate Thick-Toed Gecko	Zimbabwe, Republic of South Africa (N. Transvaal), E. Botswana, W. Mozambique (S. Blinkwater/Brak River)	Max TL 17cm: avg 12-15cm	A lighter brown gecko than most of its co-species, with pronounced tubercles on the body and short darker brown patches, or stripes, on its dorsal surfaces. The tail is fairly long and tapering. Click here to find a photograph. B I
P. turneri	Turner's Thick-Toed Gecko	RSA (Orange River Valley, Free State, NW and N Provinces, N KwaZulu-Natal), Namibia, Botswana, Zimbabwe, Angola and Tanzania; poss. Kenya		??. B I
P. vansoni	Van Son's Thick-Toed Gecko	RSA (Free State, E N Provinces, Mpumalanga, KwaZulu-Natal), SE Zimbabwe, Mozambique	8-12cm	B I
P. vanzyli	Namib Desert Gecko	S Angola, Namibia		Little is available on P. vanzyli (formerly known as Kaokogecko vanzyli). It apparently lives in those parts of the Namib Desert where gravel and rock are more prevalent than sand, which may be why the webbing on the feet is only found on the rear limbs. I have not seen these offered for sale anywhere (hardly mentioned at all, in fact), but I would imagine care to be similar as for P. rangei, with possibly more rocks and stones to simulate their native environment.
P. visseri	Visser’s Gecko	Namibia, Republic of South Africa
P. wahlbergii	Kalahari Ground Gecko, Wahlberg’s Kalahari Ground Gecko	Angola, Namibia, Botswana, Republic of South Africa	4-5"	Formerly assigned to the now invalid genus Colopus. Slender gecko with overlapping scales and somewhat reduced pads and claws. It is found on flat sandy plains with scattered vegetation (Branch), where it shelters among cover or in short burrows. There are no preanal or femoral pores, but females can be distinguished by the small hind claws. The dorsal coloration is a shade of orange or brown, and the belly is white. Diet consists of small insects.
P. w. wahlbergii	Kalahari Ground Gecko, Wahlberg’s Kalahari Ground Gecko	Kalahari desert (N Cape to Caprivi strip)		Distinguished most easily by single large pale dorsal blotches that may fuse in the centre. The head is somewhat rounded in comparison with P. w. furcifer.
P. w. furcifer	Striped Ground Gecko	W/SW Kalahari		Head is more pointed in this subspecies: there is a dark-edged pale dorsal vertebral line that divides on the neck.
P. waterbergensis		N Namibia
P. weberi	Weber's Thick-Toed Gecko	C Namibia, RSA (Namaqualand to W Cape)	7-10cm	B I
P. werneri	Werner’s Thick-Toed Gecko	Namibia

Bibliography

Field Guide to Snakes and other Reptiles of Southern Africa, Bill Branch, Struik, 1998. Best guide I have yet encountered on the members of this genus.

Field Guide to the Reptiles of East Africa by Stephen Spawls, Kim Howell, Robert Drewes and James Ashe. Detailed and invaluable review of all reptile species in the region.

Lizards of the World, Mattison

Keeping and Breeding Lizards, Mattison

Geckos: Keeping and Breeding Them in Captivity, Walls and Walls, TFH 1999.

A new species of the Pachydactylus weberi complex (Reptilia: Squamata: Gekkonidae) from the Namibrand Reserve, southern Namibia”, William R. Branch, Aaron M. Bauer, Todd R. Jackman, and Matthew Heinicke, Breviora Museum of Comparative Zoology, Number 524, Cambridge, Massachusetts, 13 June 2011. Gives details of P. etultra.

“A Revision of Pachydactylus scutatus (Reptilia: Squamata: Gekkonidae) with the Description of a New Species from Northern Namibia”, Aaron M Bauer, Trip Lamb and William R Branch, Proceedings of the California Academy of Sciences, Volume 53, No. 3, June 14 2002. Describes P. parascutatus.

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